The branch axle marks into the Botanik one of the three basic organs of the Kormophyten. It connects serving other two basic organs root and sheet to the nutrition in both directions. The branch axle carries the sheet roof and induces this as favorably as possible to its environmental condition (see plant movement). It is an organ, which developed in the course of the Landgangs of the Embryophyten. It serves stabilization, storage as well as transportation organ for the water, nutrient and Assimilatleitung.
Between the root beginning and the the Hypokotyl lies. This section of the branch is formed as the first with germinating. Between the and the beginning of the first continuation sheet the Epikotyl lies.
The branch axle is frequently somewhat thickened in the beginning places that sheets, therefore one calls this place Nodium (knot). The section between two Nodien is called dementsprechent Internodium. This Internodien is first still tossed with the Keimpflanze, whereby the sheets sitting at the Nodien sit closely one on the other. The aspect ratio of the branch axle takes place via an aspect ratio growth of the Internodien (interkalares growth). A branch axle with completely stretched Internodien is called Langtrieb, whereas a branch remains tossed short shoot is called. The interkalare education fabric, which is particularly because of the Basen of the internodien, stops the activity with a Langtrieb.
With many deciduous trees (e.g. all fruit trees) the short shoots carry the blooms and thus the fruits. Therefore they also fruit wood are called. With the larches and the Kiefern the needle sheets sit likewise on short shoots.
With some plants (e.g. Broad way Erich) is missing the aspect ratio growth of the Internodien completely, during it with others (e.g. Dachwurz) only with the bloom formation begins.
Bypasses of the branch axle result with seed plants from driving the side buds out. With some Moosen and ferns arise however also still the original dichome bypass, i.e. the vertex cells of a branch divides into two Gabelsprosse.Es gives different bypass types, from which two Grundtypen can be derived: the sympodiale bypass and the monopodiale bypass.
With the mono podium it acts around a bypass with continuous axle. By the same, akroton promoted Spitzenmeristem the before-year old impulse section is continued annually and side buds and side impulses is (e.g. Spruces and oaks).
A Sympodium is a type of bypass, with which the further growth of the rung is continued not by the centerline but by subterminalen side axles. The finalconstant bud dies thereby and the side buds drives out. (e.g. Beeches and lime trees).
If further growth is taken over of two for instance equal strong side axles, one speaks of a Dichasium (e.g. Flieder). A Monochasium is present, if only one side axle takes over further growth (e.g. Lime tree). This nearly always aligns itself thereby in the same direction as the centerline over-culminated, is exhausted then soon and again from a further side axle is over-culminated. Such Monochasium consists thus of different concatenated gradually with one another side axle and is at first sight usually hardly distinguishable by a branch with passing centerline. Thereby an illusory axle develops. A Monochasium is to be recognized by the arrangement of the sheets. Since side axles always rise from the shoulder of a sheet, the sheets at the illusory axle stand apparently for the bloom conditions opposite with a Monochasium (e.g. Common grape vine). With continuous centerline would be to be found against it the bloom conditions in the shoulders of the sheets.
Float above all the buds of the upper branch region out are this a akrotoner stature, which leads to a stature. Result the side impulses from the buds of the lower branch region are this a basitoner stature and it arise a stature.
The vegetation cone (also Apex) is the point of the branch, at which length growth carries out itself. The vegetation cone is arranged into different development zones:
The education zone is the outermost point of the cone, at which new places develop. This zone is only approx. 50 long. With the seed plants this fabric is the apical Meristem, while it is with box stems and ferns a dreischneidige vertex cell.
Behind the education zone lies the 50 to 80 long Determinationszone. Here on the differentiation each cell is decided. In the Determinationszone is already present an arrangement of the vegetation cone into a central fabric complex (corpus) and this coating Tunika. Between Korpus and Tunika a Restmeristem remains.
After the Determinationszone the aspect ratio zone follows, in which the cells differentiate themselves and stretch. The preliminary stages of the guidance bundles are formed here by the Restmeristem, which differentiates itself in this zone to a Prokambium. It forms a Protophloem outward and a Protoxylem inward. The corpus differentiates itself to parenchymatischem Mark and the Tunika to epidermis and crust. The Tunika produces also the sheet plants. In the aspect ratio zone also primary thickness growth takes place apart from aspect ratio growth.
After the differentiation of the cells are the following fabric types:
The epidermis protects the primary branch axle outward. It possesses gap openings and a Cuticula.
This fabric mostly consists of elongated cells with thickened walls. One differentiates between Sklerenchym and Kollenchym.Sklerenchym consists of dead cells and arises usually as layer around a guidance bundle. Sklerenchymzellen train thickened secondary cell walls, these are often by lignin strengthened. By the storages the cells die. They are divided in two groups:
Kollenchym is stretchable against it still growth and, not wood width unit strengthening fabric from living cells. The living cells of the Kollenchyms are usually rich at Chloroplasten, which are edges and/or individual walls by cellulose or Pektinauflagerungen strengthened.
One differentiates three different kinds from Kollenchym:
Particularly Marks consist the basic fabric in the center of the branch of Parenchym and that. Mark serves above all the storage of materials, can have torn up however with some plants, so that a Mark cave develops.
The fabric serving for transport are into strands, the guidance bundles combined. Guidance bundles are responsible for the long-distance haulage of water, solved materials, as well as organic substances (main sugars) in the branch, in the sheet and in the root of higher plants (container plants). Guidance bundles consist of the Xylem, i.e. fell trees-hurry with cell elements for the water transportation (for example tracheae and Tracheiden) and the Phloem, i.e. the phloem part, for the transport of the Assimilate with filter cells, filter tubes and escort cells.
There are different guidance bundle types: simple guidance bundles consist only of filter or fell trees-hurry. Compound guidance bundles have filter and fell trees-hurry. With the concentric guidance bundles lies seventh-hurry around fell trees-hurry (or in reverse). The most frequent type is the so-called kollaterale guidance bundle, with which seventh-hurry outside and fell trees-hurry inside lies. With open guidance bundles (comes with dikotylen plants forwards) still another Kambium steps between Xylem and Phloem in addition. In roots the guidance bundles are combined into a guidance bundle system, where fell trees-hurry like the spokes of a Rades arranged actual the phloem part lies between the spokes.
Horizontal growth is called with plants thickness growth. A primary and a secondary thickness growth can be differentiated. Primary thickness growth decreases/goes back alone to the growth already of the cells formed in the young branch of the apical Meristem (education fabric), while with secondary thickness growth by the Kambium, which lies between Phloem and Xylem, after both sides additional cells are abgegliedert, which grow into the width. Also the Korkkambium developing in the Phloem contributes to secondary thickness growth; this is particularly remarkable e.g. with the cork oak.
plants (Monokotyledonen) do not possess with few exceptions (kite trees, Yucca and club lilies) secondary separate only a primary thickness growth. Therefore palms show no taper ratio upward.
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